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The physiological
growth model BALANCE calculates the 3-dimensional development of individual
trees or forest stands and estimates the consequences of environmental
influences.
As an individual
tree model BALANCE simulates growth responses on the single tree level, which
enables also an estimation of the influence of competition, stand structure,
species mixture, and management impacts because tree development is described
as a response to individual environmental conditions and environmental conditions
change with each individual tree development. Dimensional tree growth is
calculated once a year based on the biomass increase of the woody tissue that
has been accumulated during the last year by each single tree. Biomass of an
individual tree is calculated from the dimensional variables tree position,
tree- and crown base height, diameter, and crown radii. The increase in biomass
is the result of the interaction of several physiological processes which
depend on the physical and chemical microenvironment that is itself influenced
by the stand structure.
The individual
carbon-, water- and nutrient balances of the trees species beech, oak, spruce
and pine are the fundamental processes for the simulation of growth. Each tree
is structured in crown and root layers, which are in turn divided in up to eight
crown- and root sectors. For each layer resp. each sector micro climate and
water balance are calculated by using temperature, radiation, precipitation,
humidity and wind speed measurements from climate stations. While these
calculations are computed daily, the physiological processes assimilation,
respiration, nutrient uptake, growth, senescence and allocation are calculated
in monthly or decadal time steps (= 10-day periods) from the aggregated driving
variables. This way, CO2-concentration, soil condition, competition between
individuals, and stress factors, as for example air pollution and nutrition
deficiency, can be considered besides the weather conditions when modelling the
growth of trees. Based on the individual carbon balance, dimensional changes
and mortality of a tree are computed annually.
BALANCE includes
different approaches for the estimation of the stand climate for each individual
tree. The Penman-Monteith equation is
the base of the water balance calculations. Via the stomatal closure water
balance is connected with photosynthesis, which is calculated in BALANCE by
using the approach of Haxeltine and Prentice (1996). Carbon and nitrogen allocation
into roots, branches, foliage and stem is computed according to functional
balance and pipe model principles.
To depict the
relationships between the environmental influences and growth the annual cycle
of foliage development must be known. With the beginning of bud burst foliage,
biomass and leaf area as well as light availability and radiation absorption
change. Thus, the date of foliage emergence in a tree determines its
assimilation and respiration rate but also affects the environmental conditions
of the trees in its vicinity. In BALANCE the beginning of bud burst is modelled
by using a temperature sum model (Rötzer et al. 2004), while foliage senescence
is estimated in dependence of the respiration sum.
Because tree
development is described as a response to individual environmental conditions
and environmental conditions change with the individual tree development,
environmental influences can be assessed in any kind of species mixture or
stand structure.
A more detailed
description of the model can be obtained in Grote and Pretzsch (2002), in
Rötzer et al. (2005) and in Rötzer et al. (2009a, 2009b). A review of forest
growth models and its use for forest management can be found in Pretzsch et al.
(2008).
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